I will explain the best I can.
The longer the runner length the more velocity the air charge has comming into the cumbustion chamber. This higher velocity creates a swirl effect which better mixes the fuel with the gasoline.
Furthermore, once the intake valve closes there is an area of high pressure behind the valve. This pressure bounces back through the manifold off the bottom of the plenum and back up the runner. IF the runner is long enough this area of high pressure returning to the back of the valve will occur right when the intake vavle is opening again. This crams that much more air into the C.Chamber creating a kind of turbocharging effect. This only occurs at lower rpm, and with a long enough runner.